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CC 41    Theobroma cacao

 Costa Rica - CATIE (1981)
Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.

Derivation: Cacao Center.
Collected by Paredes (& others).
Location: La Lola, Cacao Center, Costa Rica.

Notes:  Costa Rica - CATIE, 1981 [ENG81A] Selections from various populations.
 

 Parents:
UF 676  x Unknown   Costa Rica - CATIE, 1981
 Father to:
CEPEC 34    Brazil - CEPEC, 1987

 HELD IN

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List received in 2012   Reference
• Accession: 316
   - Local Name: CC 41
   - Year of Accession: 1981
List received in 1987   Reference
• Local Name: CC 41
   - Year of Accession: 1979
List received in 2018   Reference
• Accession: 10146
   - Local Name: CC-41
   - Year of Accession: 1979
List received in 2012   Reference
• Local Name: CC 41
   - Year of Accession: 1990
List received in 2012   Reference
• Local Name: CC 41
List received in 1994   Reference
• Local Name: CC 41
   - Year of Accession: 1966
 Malaysia, Agricultural Research Centre (ARC)
List received in 1990   Reference
• Local Name: CC 41
List received in 1992   Reference
• Local Name: CC 41
   - Year of Accession: 1987
List received in 2011   Reference
• Local Name: CC 41
   - Year of Accession: 1994
List received in 2016   Reference
• Accession: PI 315242     GRIN-Global 
   - Local Name: CC 41
   - Year of Accession: 1939

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Susceptible, 100.00 % Infection
Note: External Severety = 4.9 and Internal Severety = 5.0
 Phillips, W., 1997
Susceptible, 100.00 % Infection
Note: External Severety = 4.9 and Internal Severety = 5.0
 Phillips, W., 1996
Intermediate
 Morera, J. & Mora, A., 1990
Intermediate
 Enriquez, G.A. & Soria, V.J., 1967
Moderate Resistance, 85.00 % Infection, sp. P. palmivora (Pod).
Note: Unripe and not wounded
 Lopez-Baez, O. et al., 1996
Moderately Susceptible, sp. P. palmivora (Pod).
Note: Unripe, Attached and not wounded
 Morera, J., 1993
Moderately Susceptible, 100.00 % Infection, sp. P. palmivora (Pod).
Note: Unripe, Attached and not wounded
 Lawrence, J.S., 1978
Susceptible, sp. P. palmivora (Pod).
 Phillips, W., 1997
Tolerant, sp. P. palmivora (Pod).
 Enriquez, G. & Soria, V.J., 1977
Tolerant, sp. P. palmivora (Pod).
Note: (Note: Field resistance via escape in the Atlantic Zone of Costa Rica)
 Esquivel, O., 1973
90.09 % Infection
Note: 22 plants sampled..
 Subali, S., 1992
Tolerant
 Fonseca, S.E.A & Albuquerque, P.S.B. de, 1996

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Seed Index: 123.4 (dried beans per 100g)
Pod Index: 31.1 (pods per kg dried beans)
 Reference
Pod Index: 28.0 (pods per kg dried beans)
 Reference
Seed Index: 84.0 (dried beans per 100g)
Productivity: medium - high
 Reference

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Number: 39.5
Total Dry Weight: 32.1 g
Dry Weight: 0.81 g
Note: 46 plants sampled
 Reference
Colour: dark purple
Shape: ovate
Number: 43.0
Total Wet Weight: 62.0 g
Cotyledon Dry Weight: 0.83 g
Cotyledon Length: 21.7 mm
Cotyledon Width: 12.6 mm
 Reference
Colour Dark Purple: Present
Colour Light Purple: Absent
Colour Grey: Absent
Colour White: Absent
Colour Mottled: Absent
Number: 35.3
Maximum Number: 48
Cotyledon Wet Weight: 1.93 g
Cotyledon Dry Weight: 1.19 g
Cotyledon Length: 24.2 mm
Cotyledon Width: 13.7 mm
Cotyledon Thickness: 8.6 mm
 Reference
Colour Purple: 100 %
Shape: flat
Length: 25.0 mm
Width: 14.0 mm
Thickness: 9.0 mm
Cotyledon Wet Weight: 1.99 g
Cotyledon Dry Weight: 1.18 g
 Reference
Colour Dark Purple: 100 %
Cotyledon Wet Weight: 1.99 g
Cotyledon Dry Weight: 1.18 g
Cotyledon Length: 25.0 mm
Cotyledon Width: 14.0 mm
Cotyledon Thickness: 9.0 mm
 Reference
Number: 36.2
Dry Weight: 0.98 g
 Reference

Butterfat Content: 52.10 %
Maximum Butterfat Content: 54.01 %
Minimum Butterfat Content: 50.23 %
Note: 11 plants sampled
 Reference
Testa: 6.50 %
 Reference

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Shape: elliptical
Basal Constriction: strong
Apex Form: obtuse
Rugosity: slight
Length: 140.0 mm
Breadth: 79.0 mm
Colour: anthocyanin absent
 Reference
Rugosity: intense
 Reference
Basal Constriction: slight
Apex Form: obtuse
Rugosity: slight
Husk Thickness: 9.8 mm (11.7 - 7.9)
Length: 158.0 mm
Breadth: 85.0 mm
Colour: absent anthocyanin
Weight: 523.0 g
Pods Per Tree: 6
Hardness: hard
Length Breadth Ratio: 1.85
Note: ratio of distances between ridges within a pair and between ridges of two different pairs = 0.68
 Reference
Shape: amelonado
Husk Thickness: 16.8 mm
Length: 162.0 mm
Breadth: 96.0 mm
Colour: yellow
Hardness: very hard
Length Breadth Ratio: 1.75
 Reference
Shape: angoleta
Rugosity: moderate
Husk Thickness: 16.8 mm
Length: 162.0 mm
Breadth: 96.0 mm
Colour: yellow
Hardness: hard
Length Breadth Ratio: 1.68
 Reference
Shape: amelonado
Colour: green
 Reference
Length: 149.6 mm
Breadth: 83.0 mm
Weight: 429.0 g
Length Breadth Ratio: 1.80
 Reference

Accession: RUQ 1428
Light Saturated Photosynthetic Rate: 3.69 μmol m2 s-1
Stomatal Conductance: 0.06 mol m2 s-1
Transpiration Rate: 1.00 mmol m2 s-1
Intrinsic Water Use Efficiency: 61.09 μmol mol-1
Instantaneous Water Use Efficiency: 3.43 mmol mol-1
Flush Size: 4
Leaf Area: 199.90 cm2
Leaf Dry Weight: 1.25 g
Specific Leaf Area: 163.40 cm2 g-1
 Reference

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Colour: intense anthocyanin
 Reference
Colour: brownish red
 Reference
Colour: intense anthocyanin
 Reference
Colour: yellow
 Reference

Length to Width Ratio: 2.91
Basal Angle: 154.0 °
Apical Angle: 84.0 °
 Reference
Length to Width Ratio: 2.91
Basal Angle: 154.0 °
Apical Angle: 84.0 °
 Reference

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Self Compatibility: self compatible
 Reference
Ligule Colour: anthocyanin absent
Stamen Colour: moderate anthocyanin
Sepal Length: 7.53 mm
Ligule Width: 2.23 mm
Ovule Number: 42
Style Length: 2.40 mm
Pedicel Colour: red
 Reference
Self Compatibility: self compatible
 Reference
Ligule Colour: slight anthocyanin (apical)
Stamen Colour: slight anthocyanin
Sepal Colour: pale green
Sepal Width: 2.11 mm
Sepal Length: 9.35 mm
Ligule Length: 8.11 mm
Ligule Width: 2.09 mm
Ovary Colour: anthocyanin absent (apical)
Ovary Width: 0.99 mm
Ovary Length: 1.43 mm
Style Length: 2.01 mm
Staminode Length: 7.13 mm
Pedicel Colour: pastel red
Self Compatibility: self compatible
Note: Translation of Methuen colour codes.
 Reference
Bud Colour: moderate anthocyanin
Ligule Colour: slight anthocyanin
Stamen Colour: intense anthocyanin
Guide Line Length: 3.60 mm
Sepal Colour: slight anthocyanin
Sepal Width: 2.50 mm
Sepal Length: 9.20 mm
Sepal Length to Width Ratio: 3.64
Ligule Length: 4.40 mm
Ligule Width: 2.60 mm
Ligule Length to Width Ratio: 1.72
Ribbon Length: 4.20 mm
Ribbon Colour: slight anthocyanin
Ovary Colour: anthocyanin absent (apical), anthocyanin absent (basal)
Ovary Width: 1.30 mm
Ovary Length: 2.20 mm
Ovary Length to Width Ratio: 1.68
Ovule Number: 45
Style Colour: anthocyanin absent (apical), anthocyanin absent (basal)
Style Length: 2.00 mm
Staminode Colour: intense anthocyanin
Staminode Length: 8.90 mm
Pedicel Length: 18.80 mm
Pedicel Colour: intense anthocyanin, intense anthocyanin (ab), slight anthocyanin (basal)
Hairs on Upper Surface of Sepal: present
Pigmented Glands on Lower Surface of Sepal: present
Horns at Base of Guide Lines: absent
Pigmented Spot on Pollen Sac: present
 Reference
Bud Colour: green
Ligule Colour: green
Stamen Colour: moderate anthocyanin
Guide Line Length: 3.60 mm
Sepal Colour: green
Sepal Width: 2.50 mm
Sepal Length: 9.20 mm
Ligule Length: 4.40 mm
Ligule Width: 2.60 mm
Ribbon Length: 4.20 mm
Ribbon Colour: green
Ovary Width: 1.30 mm
Ovary Length: 2.20 mm
Ovule Number: 45
Style Colour: white (apical), white (basal)
Style Length: 2.00 mm
Staminode Colour: yellow
Staminode Length: 8.90 mm
Pedicel Length: 18.80 mm
Pedicel Colour: yellow, yellow (ab), green (basal)
Hairs on Upper Surface of Sepal: present
Pigmented Glands on Lower Surface of Sepal: present
Horns at Base of Guide Lines: absent
 Reference
Self Compatibility: self compatible
 Reference

Costa Rica - CATIE (1981) Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.
Brazil - CEPEC (1987) List of clones in the CEPEC, Bahia, Brazil collection. Computer printout from CEPEC. Supplied by B.G.D. Bartley, CEPEC, Brazil.
CEPEC (2012) Accession list for the CEPEC (Bahia, Brazil) collection. Dr. José Luis Pires, personal communication, February 2012.
Almeida, C.M.V.C. de, Barriga, J.P., Machado, P.F.R. & Bartley, B.G.D. (1987) Evolucao do programa de conservacao dos recursos geneticos de cacau na Amazonia Brasileira. CEPLAC/DEPEA, Boletim Tecnico, Belem, Brasil 5: pp.108.
Allan Mata (2018) Accession list for the International Cacao Collection at CATIE (IC3). Personal communication.
Philippe Lachenaud (2012) F. Guiana database, CIRAD-CP. Data supplied to ICGD by email. Dr. Philippe Lachenaud, CIRAD Montpellier, France, January 2012.
CRIG (2012) Cocoa accessions held in the Cocoa Research Institute of Ghana (CRIG). Francis Kwame Padi, personal communication, February 2012.
Guatemala - Los Brilliantes (1994) Lista de clones existentes en la Estacion de Investigacion y Produccion Agricola "Los Brillantes" a Agosto 1994. M.T. Reyes Barrios, EIAP Guatemala. Personal Communication.
Malaysia - ARC Tuaran (1990) List of clones held in the Dept. of Agriculture's collection, at Sabah, Tuaran, Malaysia. E.B. Tay, Malaysian Cocoa Board, Malaysia. Personal Communication.
Lopez-Baez, O., Mulato-Brito, J. & Lopez-Arroyo, J. (1992) Recursos geneticos del cacao, Theobroma cacao L. disponibles en Mexico. Agricultura Tecnico Mexicana 18(1-2): pp. 25-38.
CRU (2011) ICG,T accessions. CRU website.
USDA, ARS Tropical Agriculture Research Station, Mayaguez, Puerto Rico. (2016) List of cocoa accessions held at Mayaguez in April 2016. USDA-ARS GRIN-Global. [Online Database] National Germplasm Resources Laboratory, Beltsville, Maryland.
Phillips, W. (1997) Results of the artificial inoculation of Moniliophthora rorei using the Aspersion method. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Phillips, W. (1996) Cocoa cultivar reaction when innoculated with Moniliophthora roreri. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Morera, J. & Mora, A. (1990) Banco de Germoplasma de Cacao del CATIE. Information supplied on computer diskette by CATIE, Turrialba, Costa Rica.
Enriquez, G.A. & Soria, V.J. (1967) Cacao Cultivars Register. IICA Teaching and Research Centre, Costa Rica
Lopez-Baez, O., Fraire, V.G. & Cueto, M.J. (1996) Mejoramiento genetico para la resistencia a Phytophthora en Mexico. Proceedings of the International Workshop on the Contribution of Disease Resistance to Cocoa Variety Improvement, INGENIC, Salvador, 24th-26th September, 1996. pp.51-55.
Morera, J. (1993) Reaction of clones to Phytophthora palmivora. J. Morera, CATIE, Costa Rica. Personal Communication.
Lawrence, J.S. (1978) Screening of cacao cultivars for resistance to Phytophthora palmivora in the collection at CATIE, Costa Rica. Revista Theobroma 8: pp. 125-131.
Phillips, W. (1997) Results of the artificial inoculation of Phytophthora palmivora using the Paper Disc method. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Enriquez, G. & Soria, V.J. (1977) Mejoramiento genético para resistencia a cinco enfermedades del cacao. In: Proceedings of the 6th International Cocoa Research Conference, Caracas, Venezuela, 1977. Cocoa Producers' Alliance, Lagos, Nigeria. pp. 45-56.
Esquivel, O. (1973) Estudios sobre la reaccion de resistencia de algunos cultivares de cacao (Theobroma cacao L.) a la pudricion de frutos causada por Phytophthora palmivora (Butl.) Butl. MSc Thesis, IICA, Turrialba, Costa Rica.
Subali, S. (1992) MARDI's results on performance of imported primary cocoa clones. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Fonseca, S.E.A & Albuquerque, P.S.B. de (1996) Avaliacao de clones de cacau na Amazonia Brasileira em relaçao a incidencia de vassoura-de bruxa. In: Proceedings of the 12th International Cocoa Research Conference, Salvador, Brazil, 1996. Cocoa Producers' Alliance.
Malaysia - BAL Plantations (1991) Results from cocoa butter content determinations using Nuclear Magnetic Resonance (NMR) analysis. G. Lockwood, CDC, U.K. Personal Communication.
Frances Bekele & Gillian Bidaisee (2022) Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Palaniappan, S. & Shuhaimi Shamsuddin (1992) Early evaluation of some cocoa germplasm at Pusut Perkhidmatan Pertanian Tun Razak. In: Unpublished Proceedings of MCGC Mini-Seminar on the Performance of Imported Cocoa Clones, 13th October, 1992. Kuala Lumpur, Malaysia.
Morera, J., Paredes, A. & Mora, A. (1991) Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia. IICA, San Jose, Costa Rica, pp. 49.
Fiona Lahive (2017) Phenotypic data from climate change trials. Personal communication.