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RIM 101 [MEX] Group Name Theobroma cacao

Hunter, J.R. (1961)
The status of cacao variety improvement work in the Western Hemisphere. Cacao (Costa Rica) 4: pp. 1-16.

Derivation: Rosario Izapa Mexico.
Collected by del Arco & Spencer.
Location: Tuxclachico, Chiapas, Mexico.

Hunter, 1961a [HUN61A]. Fruit - Long, thin Soconusco type. Seed - white. Bartley, 1997a [BAR97A]. Most of the genotypes are very similar with regard to their vegetative and floral characteristics. The fruit shapes also show very little variation as do most other fruit characteristics. Most of these genotypes have fruit with a light green colour and the others have some degree of anthocyanin pigmentation.

Notes: Hunter, 1961 [HUN61A] Selections from Hda La Rioja. Selections from the Pacific coast with some MATINA genes.

In the original format of the name was composed of a number with "R" or "RIM" as suffix to that.

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Synonyms: R 101 [MEX]

HELD IN

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Costa Rica, Centro Agronómico Tropical de Investigación y Enseñanza (CATIE): List received in 2018 Reference

Reference

Allan Mata 2018
Accession list for the International Cacao Collection at CATIE (IC3). Personal communication.
Notes

Excel file received by email.; • Accession: 10440; - Local Name: RIM-101; - Year of Accession: 1958
France, CIRAD (Centre de Coopération Internationale en Researche Agronomique pour le Développment): List received in 2012 Reference

Reference

Philippe Lachenaud 2012
CIRAD Montpellier Clone List (Excel Spreadsheet). Data supplied to ICGD by email. Dr. Philippe Lachenaud, CIRAD Montpellier, France, January 2012.; • Local Name: RIM 101; - Year of Accession: 1993
French Guiana, CIRAD (Centre de Coopération Internationale en Researche Agronomique pour le Développment): List received in 2012 Reference

Reference

Philippe Lachenaud 2012
F. Guiana database, CIRAD-CP. Data supplied to ICGD by email. Dr. Philippe Lachenaud, CIRAD Montpellier, France, January 2012.; • Local Name: RIM 101; - Year of Accession: 1992
Guatemala, Estacion Exeparimental Los Brillantes: List received in 1994 Reference

Reference

Guatemala - Los Brilliantes 1994
Lista de clones existentes en la Estacion de Investigacion y Produccion Agricola "Los Brillantes" a Agosto 1994. M.T. Reyes Barrios, EIAP Guatemala. Personal Communication.; • Local Name: R 101 [MEX]; - Year of Accession: 1955
Honduras, Fundación Hondureña de Investigación Agrícola (FHIA): List received in 1994 Reference

Reference

Honduras - CEDEC 1994
List of accessions in CEDEC, La Masica, Honduras. J.A. Sanchez, FHIA, Honduras. Personal Communication.; • Local Name: R 101 [MEX]
Malaysia, Agricultural Research Centre (ARC): List received in 1994 Reference

Reference

Malaysia - ARC Tuaran 1994
Accessions in the Malaysian Cocoa Board collection obtained from USDA Mayaguez since 1991. M.T. Lee, MCB, Malaysia. Personal Communication.; • Local Name: R 101 [MEX]; - Year of Accession: 1992
Malaysia, Malaysian Cocoa Board (MCB): List received in 1998 Reference

Reference

Malaysia - ARC Tuaran 1998
Accessions held in the Malaysian Cocoa Board collection, May 1998. K. Lamin, MCB, Malaysia, Personal Communication.; • Local Name: R 101 [MEX]; - Year of Accession: 1993
Mexico, Campo Experimental Rosario Izapa, INIFAP: List received in 1992 Reference

Reference

Lopez-Baez, O., Mulato-Brito, J. & Lopez-Arroyo, J. 1992
Recursos geneticos del cacao, Theobroma cacao L. disponibles en Mexico. Agricultura Tecnico Mexicana 18(1-2): pp. 25-38.; • Local Name: RIM 101 [MEX]; - Year of Accession: 1945
Trinidad and Tobago, International Cocoa Genebank, Trinidad (ICG,T): List received in 2011 Reference

Reference

CRU 2011
ICG,T accessions. CRU website.
Notes

Website visited 12th April 2011

Link Unavailable [http://sta.uwi.edu/cru/icgt-accessions.asp]; • Local Name: RIM 101 [MEX]; - Year of Accession: 1994
Venezuela, MAC Caucagua: List received in 1972 Reference

Reference

Perez, A.Z., Reyes, H.E. & Reyes, L.C. 1972
Programa de mejoramiento genetico de cacao en Venezuela. MAC, Caucagua, Venezuela, pp. 131.; • Local Name: R 101 [MEX]

DISEASE

Frosty Pod Rot: Susceptible, 100.00 % Infection; Note: External Severety = 4.8 and Internal Severety = 5.0; Phillips, W., 1997

Reference

Phillips, W. 1997
Results of the artificial inoculation of Moniliophthora rorei using the Aspersion method. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Notes

Costa Rica - CATIE Datafile: DIS_MODS - Disease reaction to Monilia Pod Rot Descriptors: Resistance based on severity of infection of Moniliophthora roreri using the aspersion method Environment: CATIE, Costa Rica Germplasm: Clones held at the CATIE germplasm collection Experimental conditions: External severity of infection: scale of six values, assessed 9 weeks after inoculation (0 = healthy pod, 5 = necrosis and sporulation). Internal severity of infection: scale of six values, assessed 9 weeks after inoculation (0 = 0% of internal tissues affected, 5 = more than 80%). This parameter was used to classify the cultivars. Incidence of infection: sick pods in relation to inoculated pods.

Disease
Phytophthora: Susceptible, sp. P. palmivora (Pod).; Phillips, W., 1997

Reference

Phillips, W. 1997
Results of the artificial inoculation of Phytophthora palmivora using the Paper Disc method. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication
Notes

Costa Rica - CATIE Datafile: DIS_PHDS - Disease reaction to Phytophthora Descriptors: Resistance to Phytophthora palmivora and P. megakarya Environment: Artificial inoculation using the paper disk method. Resisance based on diameter of the lesion ten days after inoculation with 0.0015 zoopsores/ml. Resistant = 0-3 cm; moderately resistant = 3.1 - 6 cm; moderately susceptible = 6.1 - 9 cm; susceptible = > 9.1 cm lesion diameter. Germplasm: 188 Clones selected from the collection at CATIE, Costa Rica Disease - Phytophthora palmivora, P. megakarya (Local Names: Black Pod, Podredumbre negra, Pourriture Brune, Podridao parda).

Disease

YIELD

Frances Bekele & Gillian Bidaisee, 2022: Pod Index: 32.3 (pods per kg dried beans); Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Lopez-Baez, O., 1995: Seed Index: 90.0 (dried beans per 100g); Pod Index: 21.9 (pods per kg dried beans); Reference

Reference

Lopez-Baez, O. 1995
Caracteristicas del fruto del germoplasma de cacao, Theobroma cacao L., seleccionado en Rosario Izapa, Chiapas. Agricultura Tecnico Mexicana 21(2): pp. 127-137.
Notes

Mexico - Rosario Izapa, Chiapas Datafile: MEX___DS MEX - Description of RIM [MEX] clones Environment: No details. Descriptors: Fruit characteristics are recorded. See also associated paper on flower and seed characteristics - Lopez-Baez & Mendez-Wong 1992 [LOP92A].

Yield Bean Butterfat Fruit

BEAN

Frances Bekele & Gillian Bidaisee, 2022: Colour: dark purple; Shape: oblong; Number: 31.0; Total Wet Weight: 52.0 g; Cotyledon Dry Weight: 1.00 g; Cotyledon Length: 21.7 mm; Cotyledon Width: 12.4 mm; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Lopez-Baez, O., 1995: Number: 29.6; Length: 23.7 mm; Width: 13.5 mm; Thickness: 8.9 mm; Dry Weight: 1.20 g; Cotyledon Dry Weight: 1.11 g; Reference

Reference

Lopez-Baez, O. 1995
Caracteristicas del fruto del germoplasma de cacao, Theobroma cacao L., seleccionado en Rosario Izapa, Chiapas. Agricultura Tecnico Mexicana 21(2): pp. 127-137.
Notes

Mexico - Rosario Izapa, Chiapas Datafile: MEX___DS MEX - Description of RIM [MEX] clones Environment: No details. Descriptors: Fruit characteristics are recorded. See also associated paper on flower and seed characteristics - Lopez-Baez & Mendez-Wong 1992 [LOP92A].

Yield Bean Butterfat Fruit

BUTTERFAT

Frances Bekele & Gillian Bidaisee, 2022: Testa: 5.40 %; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Lopez-Baez, O., 1995: Testa: 7.50 %; Reference

Reference

Lopez-Baez, O. 1995
Caracteristicas del fruto del germoplasma de cacao, Theobroma cacao L., seleccionado en Rosario Izapa, Chiapas. Agricultura Tecnico Mexicana 21(2): pp. 127-137.
Notes

Mexico - Rosario Izapa, Chiapas Datafile: MEX___DS MEX - Description of RIM [MEX] clones Environment: No details. Descriptors: Fruit characteristics are recorded. See also associated paper on flower and seed characteristics - Lopez-Baez & Mendez-Wong 1992 [LOP92A].

Yield Bean Butterfat Fruit

FRUIT

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Frances Bekele & Gillian Bidaisee, 2022: Shape: elliptical; Basal Constriction: slight; Apex Form: acute; Rugosity: moderate; Length: 156.0 mm; Breadth: 76.0 mm; Colour: slight anthocyanin; Hardness: moderate; Note: Hardness > 1.6 and <= 2.0 Mpa; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Costa Rica - CATIE, 1981: Basal Constriction: very slight; Apex Form: acute; Rugosity: moderate; Colour: moderate / slight - moderate anthocyanin (ridge/furrow); Reference

Reference

Costa Rica - CATIE 1981
Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.
Notes

Costa Rica - CATIE Datafile: CATIE__DS CRI - CATIE 1 IMAGE_DS PROGENY_ Germplasm: Descriptor information is presented for 294 clones, 8 related species and on one hybrid. In addition, limited information on a further 119 clones is given. This germplasm is mainly of Latin American origin, including the United Fruit clones, UF, and genotypes from Mexico and Brazil. Environment: Each clone is represented in the collection by up to 10 trees planted in a single row. Spacing is usually 2m within the row and 3m between the rows, although parts of the collection are planted in a 2.2 x 2.2m grid. Fertilizer, N-P-K-Mg/Ca in the ratio 18:10:6:5, is applied four times a year at 150g per tree. Shade is provided in most of the collection by an almost complete closed cover of trees of Erythrina poepiggiana and Inga spp. spaced at 12 x 12m. Character data: Seed dry weight: sample is dried for 1.5 h at 130°C and cooled in a desiccator. Seed length, seed width, seed thickness: average of 5 seeds from 20 fruits. Fruit, wall thickness at ridge: fruits are transversally sectioned and the soft endocarp tissue removed. Fruit, ridge pair separation: mean ratio of the distance between ridges within a pair and the distances between the ridges of two different pairs. Leaf, flush colour: measured when leaves are 5cm long. Producitvity: values are given as: 1=very low - c. 40 pods harvested over 3 years; 3=low, 5=intermediate - c. 40 pods obtained from 3-4 harvests over 1 year; 7=high, 9=very high - c. 40 pods harvested at one time. Pod outline drawing for each accession.

Yield Bean Fruit Flush Flower
Lopez-Baez, O., 1995: Shape: cundeamor; Rugosity: intense; Husk Thickness: 14.5 mm (12 - 17); Length: 196.0 mm; Breadth: 94.0 mm; Colour: yellow; Weight: 630.2 g; Husk Weight: 506.10 g; Length Breadth Ratio: 2.08; Reference

Reference

Lopez-Baez, O. 1995
Caracteristicas del fruto del germoplasma de cacao, Theobroma cacao L., seleccionado en Rosario Izapa, Chiapas. Agricultura Tecnico Mexicana 21(2): pp. 127-137.
Notes

Mexico - Rosario Izapa, Chiapas Datafile: MEX___DS MEX - Description of RIM [MEX] clones Environment: No details. Descriptors: Fruit characteristics are recorded. See also associated paper on flower and seed characteristics - Lopez-Baez & Mendez-Wong 1992 [LOP92A].

Yield Bean Butterfat Fruit
Morera, J. et al., 1991: Shape: cundeamor; Colour: green; Reference

Reference

Morera, J., Paredes, A. & Mora, A. 1991
Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia. IICA, San Jose, Costa Rica, pp. 49.
Notes

Costa Rica - CATIE Datafile: DIS_MODS - Disease reaction to Monilia Pod Rot CATIE3DS CRI - CATIE 3 DIS_PHDS - Disease reaction to Phytophthora DIS_CEDS - Disease reaction to Ceratocystis PROGENY_ Germplasm: Descriptions are given for 559 clones. More information on 44 of them is given in the Table CATIE, Costa Rica (1). Environment: Details are given in the Table CATIE, Costa Rica (1). Descriptors: Population type, self-compatibility, resistance to disease, pod shape and colour are described. Population types - CATIE, Costa Rica (3). Disease - Phytophthora palmivora, P. megakarya (Local Names: Black Pod, Podredumbre negra, Pourriture Brune, Podridao parda), Moniliophthora / Monilia roreri (local names: Watery Pod Rot, Moniliasis, Moniliose), Ceratocystis fimbriata (local names: Ceratocystis Wilt, Mal de Machete, Mort Subite).

Disease Fruit Flower

FLUSH

Frances Bekele & Gillian Bidaisee, 2022: Colour: moderate anthocyanin; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Costa Rica - CATIE, 1981: Colour: olive brown; Reference

Reference

Costa Rica - CATIE 1981
Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.
Notes

Costa Rica - CATIE Datafile: CATIE__DS CRI - CATIE 1 IMAGE_DS PROGENY_ Germplasm: Descriptor information is presented for 294 clones, 8 related species and on one hybrid. In addition, limited information on a further 119 clones is given. This germplasm is mainly of Latin American origin, including the United Fruit clones, UF, and genotypes from Mexico and Brazil. Environment: Each clone is represented in the collection by up to 10 trees planted in a single row. Spacing is usually 2m within the row and 3m between the rows, although parts of the collection are planted in a 2.2 x 2.2m grid. Fertilizer, N-P-K-Mg/Ca in the ratio 18:10:6:5, is applied four times a year at 150g per tree. Shade is provided in most of the collection by an almost complete closed cover of trees of Erythrina poepiggiana and Inga spp. spaced at 12 x 12m. Character data: Seed dry weight: sample is dried for 1.5 h at 130°C and cooled in a desiccator. Seed length, seed width, seed thickness: average of 5 seeds from 20 fruits. Fruit, wall thickness at ridge: fruits are transversally sectioned and the soft endocarp tissue removed. Fruit, ridge pair separation: mean ratio of the distance between ridges within a pair and the distances between the ridges of two different pairs. Leaf, flush colour: measured when leaves are 5cm long. Producitvity: values are given as: 1=very low - c. 40 pods harvested over 3 years; 3=low, 5=intermediate - c. 40 pods obtained from 3-4 harvests over 1 year; 7=high, 9=very high - c. 40 pods harvested at one time. Pod outline drawing for each accession.

Yield Bean Fruit Flush Flower

FLOWER

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Frances Bekele & Gillian Bidaisee, 2022: Ligule Colour: anthocyanin absent; Stamen Colour: anthocyanin absent; Sepal Length: 7.54 mm; Ligule Width: 2.63 mm; Ovule Number: 37; Style Length: 2.20 mm; Pedicel Colour: red; Reference

Reference

Frances Bekele & Gillian Bidaisee 2022
Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.
Notes

This single reference supercedes data entered previously and will be kept up-to-date to allow this information to be grouped together.

Yield Bean Butterfat Fruit Flush Flower
Lopez-Baez, O. & Mendel-Wong, D., 1992: Sepal Width: 2.60 mm; Sepal Length: 8.80 mm; Ligule Length: 3.90 mm; Ligule Width: 3.20 mm; Ovary Width: 1.00 mm; Ovary Length: 2.00 mm; Ovule Number: 42; Style Length: 2.50 mm; Staminode Length: 7.50 mm; Pedicel Length: 2.20 mm; Reference

Reference

Lopez-Baez, O. & Mendel-Wong, D. 1992
Caracteristicas de la flor y semilla del germoplasma de cacao, Theobroma cacao L., seleccionado en Rosario Izapa, Chiapas. Agricultura Tecnico Mexicana 18(1-2): pp. 39-48.
Notes

Morphological characters. Datafile: MEX___DS MEX - Description of RIM [MEX] clones Environment: No details. Descriptors: Flower and seed characteristics are recorded. See also associated paper on fruit characteristics - Lopez-Baez (1995) [LOP95B].

Flower
Morera, J. et al., 1991: Self Compatibility: self incompatible; Reference

Reference

Morera, J., Paredes, A. & Mora, A. 1991
Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia. IICA, San Jose, Costa Rica, pp. 49.
Notes

Costa Rica - CATIE Datafile: DIS_MODS - Disease reaction to Monilia Pod Rot CATIE3DS CRI - CATIE 3 DIS_PHDS - Disease reaction to Phytophthora DIS_CEDS - Disease reaction to Ceratocystis PROGENY_ Germplasm: Descriptions are given for 559 clones. More information on 44 of them is given in the Table CATIE, Costa Rica (1). Environment: Details are given in the Table CATIE, Costa Rica (1). Descriptors: Population type, self-compatibility, resistance to disease, pod shape and colour are described. Population types - CATIE, Costa Rica (3). Disease - Phytophthora palmivora, P. megakarya (Local Names: Black Pod, Podredumbre negra, Pourriture Brune, Podridao parda), Moniliophthora / Monilia roreri (local names: Watery Pod Rot, Moniliasis, Moniliose), Ceratocystis fimbriata (local names: Ceratocystis Wilt, Mal de Machete, Mort Subite).

Disease Fruit Flower

REFERENCES

Hunter, J.R. (1961) The status of cacao variety improvement work in the Western Hemisphere. Cacao (Costa Rica) 4: pp. 1-16.

Allan Mata (2018) Accession list for the International Cacao Collection at CATIE (IC3). Personal communication.

Philippe Lachenaud (2012) CIRAD Montpellier Clone List (Excel Spreadsheet). Data supplied to ICGD by email. Dr. Philippe Lachenaud, CIRAD Montpellier, France, January 2012.

Philippe Lachenaud (2012) F. Guiana database, CIRAD-CP. Data supplied to ICGD by email. Dr. Philippe Lachenaud, CIRAD Montpellier, France, January 2012.

Guatemala - Los Brilliantes (1994) Lista de clones existentes en la Estacion de Investigacion y Produccion Agricola "Los Brillantes" a Agosto 1994. M.T. Reyes Barrios, EIAP Guatemala. Personal Communication.

Honduras - CEDEC (1994) List of accessions in CEDEC, La Masica, Honduras. J.A. Sanchez, FHIA, Honduras. Personal Communication.

Malaysia - ARC Tuaran (1994) Accessions in the Malaysian Cocoa Board collection obtained from USDA Mayaguez since 1991. M.T. Lee, MCB, Malaysia. Personal Communication.

Malaysia - ARC Tuaran (1998) Accessions held in the Malaysian Cocoa Board collection, May 1998. K. Lamin, MCB, Malaysia, Personal Communication.

Lopez-Baez, O., Mulato-Brito, J. & Lopez-Arroyo, J. (1992) Recursos geneticos del cacao, Theobroma cacao L. disponibles en Mexico. Agricultura Tecnico Mexicana 18(1-2): pp. 25-38.

CRU (2011) ICG,T accessions. CRU website.

Perez, A.Z., Reyes, H.E. & Reyes, L.C. (1972) Programa de mejoramiento genetico de cacao en Venezuela. MAC, Caucagua, Venezuela, pp. 131.

Phillips, W. (1997) Results of the artificial inoculation of Moniliophthora rorei using the Aspersion method. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication

Phillips, W. (1997) Results of the artificial inoculation of Phytophthora palmivora using the Paper Disc method. Unpublished data supplied by W. Phillips, CATIE, Costa Rica. Personal communication

Frances Bekele & Gillian Bidaisee (2022) Morphological data from the International Cocoa Collection (ICG,T) maintained by the Cocoa Research Centre (CRC), Trinidad & Tobago. Unpublished data on fruit, bean, flower and flush morphology supplied as an Excel spreadsheet by Frances Bekele. Last update March 2022.

Lopez-Baez, O. (1995) Caracteristicas del fruto del germoplasma de cacao, Theobroma cacao L., seleccionado en Rosario Izapa, Chiapas. Agricultura Tecnico Mexicana 21(2): pp. 127-137.

Costa Rica - CATIE (1981) Genetic Resources of Cacao. A Catalogue of the CATIE Collection. J.M.M. Engels, Tropical Agricultural Research and Training Centre, CATIE, Turrialba, Costa Rica. Technical Bulletin No. 7: pp. 191.

Morera, J., Paredes, A. & Mora, A. (1991) Germoplasma de cacao en el CATIE entre 1947 y 1991, Programa II: Generacion y transferencia de tecnologia. IICA, San Jose, Costa Rica, pp. 49.

Lopez-Baez, O. & Mendel-Wong, D. (1992) Caracteristicas de la flor y semilla del germoplasma de cacao, Theobroma cacao L., seleccionado en Rosario Izapa, Chiapas. Agricultura Tecnico Mexicana 18(1-2): pp. 39-48.